Opportunity Information: Apply for PD 04 1127

  • The National Science Foundation in the science and technology and other research and development sector is offering a public funding opportunity titled "Population and Evolutionary Processes" and is now available to receive applicants.
  • Interested and eligible applicants and submit their applications by referencing the CFDA number(s): 47.074 Biological Sciences.
  • This funding opportunity was created on May 27, 2009 and posted on Aug 15, 2006.
  • Applicants must submit their applications by archived replaced by 09 1127. (Agency may still review applications by suitable applicants for the remaining/unused allocated funding in 2026.)
  • Eligible applicants include: Unrestricted (i.e., open to any type of entity above), subject to any clarification in text field entitled Additional Information on Eligibility.
Apply for PD 04 1127

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Opportunity Summary:

The National Science Foundation (NSF) Population and Evolutionary Processes opportunity (PD 04-1127) is a discretionary research grant program in the Biological Sciences (CFDA 47.074) that supports studies aimed at explaining why populations and species vary, how that variation is maintained or changes over time, and how small-scale evolutionary processes can scale up to larger macroevolutionary patterns. At its core, the program is interested in population-level properties and evolutionary mechanisms that shape diversity within populations, among populations, and among species. The scope explicitly includes both empirical and theoretical work, and it welcomes a wide range of methodological approaches, including analytical modeling, stochastic frameworks, and statistical inference.

Scientifically, the opportunity emphasizes research on microevolution and adaptation, geographic differentiation, natural hybridization, and speciation, along with broader processes that generate macroevolutionary patterns in trait evolution. A key boundary is that proposals centered on interactions among species (for example, community-level dynamics, interspecific competition, or multi-species ecological networks) are expected to be directed to NSF's Ecological Biology Cluster instead. In contrast, this cluster is a better fit for studies that are primarily organism- or lineage-focused and that use populations as the main unit for understanding evolutionary change and diversification.

Within the Population and Evolutionary Processes Program itself, NSF highlights several major topic areas. One major track is population demography, especially research on age-structured and stage-structured populations and how populations change through time. This can include work that develops or applies demographic models and uses analytical, stochastic, or statistical approaches to estimate vital rates, forecast population trajectories, or test hypotheses about population regulation and change. Another strong emphasis is evolutionary ecology of single species, including life history and life cycle phenomena in terrestrial, freshwater, and wetland organisms. This includes studies of natural selection and sexual selection, and research on the causes and consequences of reproductive isolation, phylogeography, and evolution within a single lineage (sometimes framed as phyletic evolution).

The program also places significant weight on evolutionary genetics. It supports population genetics and quantitative genetics, including research that links microevolutionary mechanisms to macroevolutionary outcomes. Notably, the description explicitly includes epigenetics and development when they are used to explain the evolution of complex phenotypes, indicating an openness to modern views of inheritance and phenotype formation as long as the focus remains on evolutionary processes. In molecular population biology, the program supports studies of the sources and consequences of variation and evolutionary change at biochemical and sequence levels, including RNA and DNA sequence evolution and mobile genetic elements. It also covers research on how gene properties (such as gene number, genomic arrangement, and patterns of interaction) shape evolutionary processes, with explicit interest in the evolution of genetic architecture and evolutionary genomics.

From a proposal and program mechanics perspective, the opportunity supports awards not only through the core Population and Evolutionary Processes Program but also through related NSF mechanisms such as CAREER, OPUS, and LTREB, which can be relevant depending on an applicant's career stage and the temporal scale of the research. Unsolicited proposals are expected to follow the NSF Grant Proposal Guide (GPG). The program encourages inter- and multidisciplinary projects that cross traditional boundaries and notes a history of co-reviewing such proposals with other NSF programs, which can be important for projects spanning genetics, evolution, demography, and related fields.

The eligibility category is described as unrestricted (open to any type of entity, subject to any clarifications in the full text). The original target dates listed for full proposals in this archived announcement were January 9, 2007 and July 9, 2007, and the notice indicates the opportunity has been archived and replaced (archived May 27, 2009). Even so, the description provides a clear picture of the scientific fit: projects should be centered on population and evolutionary processes, while proposals that primarily analyze organism-centered physiology, morphology, behavior, or development should be routed to programs in the Division of Integrative Organismal Biology (IOB). Likewise, studies focused on the population ecology of marine organisms are expected to go to the Biological Oceanography Program in the Division of Ocean Sciences, reflecting NSF's division of responsibility by environment and disciplinary focus.

FAQs: NSF Population and Evolutionary Processes (PD 04-1127)

What is the NSF Population and Evolutionary Processes opportunity (PD 04-1127)?

It is a discretionary research grant program in NSF Biological Sciences (CFDA 47.074) that supports studies explaining why populations and species vary, how variation is maintained or changes over time, and how microevolutionary processes can scale to macroevolutionary patterns.

What is the program trying to understand at a high level?

The core focus is population-level properties and evolutionary mechanisms that shape diversity within populations, among populations, and among species, including how small-scale evolutionary processes can produce larger-scale patterns in trait evolution and diversification.

What kinds of research approaches does the program allow?

The scope explicitly includes both empirical and theoretical work. It welcomes a wide range of methodological approaches, including analytical modeling, stochastic frameworks, and statistical inference.

What scientific topics are emphasized by this program?

Highlighted themes include microevolution and adaptation, geographic differentiation, natural hybridization, speciation, and broader processes that generate macroevolutionary patterns in trait evolution.

What is the main unit of analysis NSF expects for a good fit?

The program is best aligned with studies that are organism- or lineage-focused and use populations as the main unit for understanding evolutionary change and diversification.

Does the program support population demography research?

Yes. A major track is population demography, especially age-structured and stage-structured populations and how populations change through time, including development or application of demographic models and estimation of vital rates and population trajectories.

What methods are called out for demographic work?

The description specifically notes analytical, stochastic, and statistical approaches to estimate vital rates, forecast population trajectories, and test hypotheses about population regulation and change.

Does the program support evolutionary ecology studies?

Yes. It emphasizes evolutionary ecology of single species, including life history and life cycle phenomena in terrestrial, freshwater, and wetland organisms.

Are natural selection and sexual selection within scope?

Yes. The opportunity explicitly includes studies of natural selection and sexual selection within its evolutionary ecology emphasis.

Are speciation and reproductive isolation topics supported?

Yes. The program highlights research on the causes and consequences of reproductive isolation, natural hybridization, and speciation.

Is phylogeography an appropriate topic area?

Yes. Phylogeography is specifically mentioned as part of the evolutionary ecology of single species and lineage-focused evolution.

Does the program support evolutionary genetics work?

Yes. Evolutionary genetics is a significant emphasis area, including population genetics and quantitative genetics, and research linking microevolutionary mechanisms to macroevolutionary outcomes.

Is epigenetics allowed under this program?

Yes, when epigenetics (and development) is used to explain the evolution of complex phenotypes and the focus remains on evolutionary processes.

Does the program include molecular population biology and sequence evolution?

Yes. It supports research on sources and consequences of variation and evolutionary change at biochemical and sequence levels, including RNA and DNA sequence evolution and mobile genetic elements.

Does the program support evolutionary genomics and genetic architecture topics?

Yes. The description includes interest in how gene properties (gene number, genomic arrangement, and patterns of interaction) shape evolutionary processes, and it explicitly notes the evolution of genetic architecture and evolutionary genomics.

What kinds of projects are not a good fit for this program?

Projects centered on interactions among species (such as community-level dynamics, interspecific competition, or multi-species ecological networks) are expected to be directed to NSF's Ecological Biology Cluster instead.

If my proposal focuses on community or multi-species interactions, where should it go?

The announcement indicates proposals primarily about interactions among species should be directed to NSF's Ecological Biology Cluster.

If my proposal is mainly about physiology, morphology, behavior, or development, is this the right program?

The description says proposals that primarily analyze organism-centered physiology, morphology, behavior, or development should be routed to programs in the Division of Integrative Organismal Biology (IOB), rather than this Population and Evolutionary Processes cluster.

What if my study is about the population ecology of marine organisms?

The notice indicates studies focused on the population ecology of marine organisms are expected to go to the Biological Oceanography Program in the Division of Ocean Sciences.

Does NSF encourage interdisciplinary proposals for this opportunity?

Yes. The program encourages inter- and multidisciplinary projects that cross traditional boundaries and notes a history of co-reviewing such proposals with other NSF programs.

Can proposals be co-reviewed with other NSF programs?

Yes. The description explicitly notes co-reviewing with other NSF programs, which can be important for projects spanning genetics, evolution, demography, and related fields.

What proposal guidance applies to submissions?

Unsolicited proposals are expected to follow the NSF Grant Proposal Guide (GPG).

Are there specific NSF award mechanisms connected to this program?

Yes. The opportunity notes awards may be supported through the core program as well as related NSF mechanisms such as CAREER, OPUS, and LTREB, depending on career stage and the temporal scale of the research.

Who is eligible to apply?

The eligibility category is described as unrestricted (open to any type of entity), subject to any clarifications in the full text.

Is this opportunity still active?

No. The announcement indicates the opportunity has been archived and replaced, with an archive date of May 27, 2009.

What were the original proposal target dates listed in the archived announcement?

The archived notice lists January 9, 2007 and July 9, 2007 as the original target dates for full proposals.

Even though it is archived, what does the description still help clarify?

It still provides clear guidance on scientific fit: proposals should be centered on population and evolutionary processes, while work primarily focused on interspecific interactions, organism-centered integrative biology topics, or marine population ecology is expected to be directed to other NSF clusters or programs as described.

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